{"id":269,"date":"2017-04-18T03:37:48","date_gmt":"2017-04-18T03:37:48","guid":{"rendered":"http:\/\/sites.warnercnr.colostate.edu\/gwhite\/?page_id=269"},"modified":"2017-04-18T03:37:48","modified_gmt":"2017-04-18T03:37:48","slug":"link-barker-jolly-seber-model","status":"publish","type":"page","link":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/link-barker-jolly-seber-model\/","title":{"rendered":"Link-Barker Jolly-Seber Model"},"content":{"rendered":"<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"><a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/index.html\">Contents<\/a> &#8211; <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/idx.htm\">Index<\/a><\/span><\/p>\n<hr \/>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: medium\"><b>Link-Barker Jolly-Seber Model<\/b><\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"><a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/pertinentliterature.htm\">Link and Barker (2005)<\/a> reparameterized the <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/recruitment_parameters.htm\">recruitment likelihood<\/a> of <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/pertinentliterature.htm\">Pradel (1996)<\/a>, ending up at a starting point of the likeliehood from <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/pertinentliterature.htm\">Schwarz and Arnason (1996)<\/a>, used in the <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/popan_model.htm\">POPAN<\/a> data type.\u00a0 Basically, the probability of entry parameters of Schwarz and Arnason (1996) are translated to a recruitment (births and immigration) parameter (<i>f<\/i>).\u00a0 The Link-Barker model conditions on the total number of animals ever caught and so the super-population size (<i>N<\/i>) and the probability that an animal has entered the population prior to the experiment [beta(0)] are no longer required in the likelihood.\u00a0 This obviates the need for the identifiability constraint on <i>p<\/i>(1) and <i>p<\/i>(<i>t<\/i>) needed for the Schwarz and Arnason (1996) <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/popan_model.htm\">POPAN<\/a> model.\u00a0 If there are no losses on capture the Link-Barker likelihood and the Pradel (1996) likelihood are equivalent.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">The Link-Barker data type has 3 <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/parameter_matrices.htm\">PIMs<\/a> for each <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/group_names.htm\">group<\/a>: phi, <i>p<\/i>, and <i>f<\/i>.\u00a0 For <i>t<\/i> occasions, the phi PIM has <i>t<\/i> &#8211; 1 parameters, the <i>p<\/i> PIM has <i>t<\/i> parameters, and the <i>f<\/i> PIM has <i>t<\/i> &#8211; 1 parameters.\u00a0 The <i>f<\/i> parameters need not be constrained to the [0, 1] interval, so you may want to use a log or identity <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/link_functions.htm\">link function<\/a> for these parameters if estimates exceed 1.\u00a0 The default link for <i>f<\/i> is the <\/span><span style=\"font-family: Arial, helvetica, sans-serif;font-size: small\"><a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/link_functions.htm\">log link<\/a><\/span><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">, regardless of what the link type is for phi and <i>p<\/i>.\u00a0 That is, unless a parameter-specific link is used, <i>f<\/i> will be computed with the log link.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Two main advantages of this reparameterized model are that the <i>f<\/i> parameters are biologically interpretable as the birth rates (actually recruitment to the population from either immigration or births) to predict the number of new animals in the population, and the super-population size (<i>N<\/i>) is no longer required in the likelihood.\u00a0 Historically, the weakest feature of the Jolly-Seber model has been the estimates of the population sizes and births for each occasion, because these parameters are based on untestable assumptions, i.e., that the unmarked animals have exactly the same capture probabilities as the marked animals.\u00a0 Any behavioral response to initial capture will violate this assumption.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Use of individual covariates in the Link-Barker model is allowed, but the meaning of the estimates can be difficult to interpret biologically.\u00a0 That is, lambda = phi + <i>f<\/i>.\u00a0 If phi and\/or <i>f<\/i> are modeled as functions of individual covariates, then the population parameter lambda is also a function of these individual covariates.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Competing models in MARK for this data type are the Pradel recruitment (<i>f<\/i>) model, Burnham&#8217;s Jolly-Seber model (which often suffers from optimization problems) and the <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/popan_model.htm\">POPAN<\/a> data type.\u00a0 Burnham&#8217;s Jolly-Seber model provides an estimate of the population size on the first occasion, and the rate of population change (lambda) for each succeeding interval.\u00a0 The <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/popan_model.htm\">POPAN<\/a> data type does provide estimates of population sizes and births (<i>B<\/i>) by occasion as derived parameters.\u00a0 None of these models have the same likelihood, so <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/qaicc.htm\">AIC<\/a> values are not comparable.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Both the <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/recruitment_parameters.htm\">Pradel recruitment<\/a> and the Link-Barker model have the <i>f<\/i> parameter.\u00a0 However, these parameters are not exactly equivalent unless there are no losses on capture.\u00a0 The main advantage of the Link-Barker model over the Pradel parameterization is that the distribution for losses on capture factors out of the likelihood.\u00a0 Unless you have many losses on capture the parameter estimates under the two parameterisations will not differ greatly.\u00a0 If you do have a large number of losses on capture then the experiment is probably of questionable value as the removal of animals may cause a change in the demographics of the study population.\u00a0 Note that the Pradel recruitment (<i>f<\/i>) model as parameterized in MARK, does not handle the losses on capture correctly, and does not incorporate the losses on capture parameter described in <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/pertinentliterature.htm\">Pradel&#8217;s (1996)<\/a> paper.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">The Link-Barker parametrization has been extended by incorporating the Pledger <\/span><span style=\"font-family: Arial, helvetica, sans-serif;font-size: small\"><a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/mixtures.htm\">mixture<\/a><\/span><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"> model on <i>p<\/i>, allowing for <\/span><span style=\"font-family: Arial, helvetica, sans-serif;font-size: small\"><a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/heterogeneity_open_models.htm\">heterogeneity<\/a><\/span><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"> of capture probabilities.<\/span><\/p>\n","protected":false},"excerpt":{"rendered":"<p>Contents &#8211; Index Link-Barker Jolly-Seber Model Link and Barker (2005) reparameterized the recruitment likelihood of Pradel (1996), ending up at a starting point of the likeliehood from Schwarz and Arnason (1996), used in the POPAN data type.\u00a0 Basically, the probability &hellip;<\/p>\n<p class=\"read-more\"> <a class=\"more-link\" href=\"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/link-barker-jolly-seber-model\/\"> <span class=\"screen-reader-text\">Link-Barker Jolly-Seber Model<\/span> Read More &raquo;<\/a><\/p>\n","protected":false},"author":117,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-269","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages\/269","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/users\/117"}],"replies":[{"embeddable":true,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/comments?post=269"}],"version-history":[{"count":1,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages\/269\/revisions"}],"predecessor-version":[{"id":270,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages\/269\/revisions\/270"}],"wp:attachment":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/media?parent=269"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}