{"id":280,"date":"2017-04-18T03:43:30","date_gmt":"2017-04-18T03:43:30","guid":{"rendered":"http:\/\/sites.warnercnr.colostate.edu\/gwhite\/?page_id=280"},"modified":"2017-04-18T03:43:30","modified_gmt":"2017-04-18T03:43:30","slug":"occupancy-estimation-multiple-states-single-season","status":"publish","type":"page","link":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/occupancy-estimation-multiple-states-single-season\/","title":{"rendered":"Occupancy Estimation Multiple States Single Season"},"content":{"rendered":"<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"><a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/index.html\">Contents<\/a> &#8211; <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/idx.htm\">Index<\/a><\/span><\/p>\n<hr \/>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: medium\"><b>Occupancy Estimation Multiple States Single Season<\/b><\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Estimation of <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/occupancy_estimation.htm\">occupancy rates<\/a> generally assume that a site or plot is occupied or not occupied.\u00a0 This model extends this concept to not occupied, and 2 levels of occupancy.\u00a0 For example, a plot may be occupied by a pair of adults that either reproduce, or do not reproduce.\u00a0 Given that the occupancy states are hierarchial, a &#8220;1&#8221; is coded for the lower level of occupancy (e.g., adults, but no young), and a &#8220;2&#8221; is coded for the higher level of occupancy (adults plus young).\u00a0 <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/pertinentliterature.htm\">Nichols et al. (2007)<\/a> developed this model with an example of California spotted owls (<i>Strix occidentalis occidentalis<\/i>) where sites were occupied by 1 or more adults, and or occupied by both adults and young.\u00a0 I will use their example to explain the 5 parameters of this model.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">The <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/real_parameter.htm\">real parameters<\/a> of this model are:<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">psi1(<i>i<\/i>) = probability that a site <i>i<\/i> is occupied regardless of reproductive state, Pr(true state = 1 or 2);<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">psi2(<i>i<\/i>) = probability that young occurred, given that the site <i>i<\/i> is occupied, Pr(true state = 2 | true state = 1 or 2);<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"><i>p<\/i>1(<i>i, t<\/i>) = probability that occupancy is detected for site <i>i<\/i>, period <i>t<\/i>, given that true state = 1, Pr(detection | true state = 1);<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\"><i>p<\/i>2(<i>i, t<\/i>) = probability that occupancy is detected for site <i>i<\/i>, period <i>t<\/i>, given that true state = 2, Pr(detection | true state = 2);<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">delta(<i>i, t<\/i>) = probability that evidence of successful reproduction is found, given detection of occupancy at site <i>i<\/i>, period <i>t<\/i>, with successful reproduction, Pr(classified state 2|true state = 2).<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Thus, psi1 and psi2 are single parameters relating to a site, whereas <i>p<\/i>1, <i>p<\/i>2, and delta are time-specific parameters, and thus the number of values in the <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/parameter_matrices.htm\">PIM<\/a> for each of these parameters equals the number of occasions.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">The product psi1*psi2 is the unconditional probability that a site is occupied by successful breeders, and is provided as a <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/derived_parameters.htm\">derived parameter<\/a> in MARK.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">The <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/encounter_histories_file.htm\">encounter histories file<\/a> for this data type consists of only the L columns, and the example from Nichols et al. (2007) is shown here.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">\/* occasions=5, groups=1 *\/<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1.221\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">011..\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1.1..\u00a0 2;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.1212\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1111.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1022.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">111.1\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0.20.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">11100\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.11..\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.011.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0110.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">11111\u00a0 3;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.0.11\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1100.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">101..\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">&#8230;0.\u00a0 2;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.111.\u00a0 2;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0.0.0\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0.210\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">01110\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0.0.1\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">&#8230;1.\u00a0 2;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0&#8230;.\u00a0 3;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1.2..\u00a0 3;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.1101\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.12..\u00a0 2;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1122.\u00a0 3;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0120.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.1111\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">11202\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">11220\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">10110\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">.10..\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">1.200\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">00&#8230;\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0122.\u00a0 2;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0.11.\u00a0 1;\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0.100\u00a0 1;<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">0..11\u00a0 1;<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">\u00a0\u00a0<\/span><br \/>\n<span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">Notice that &#8220;.&#8221; are used to indicate that a site was not surveyed on a particular occasion, and &#8220;0&#8221;, &#8220;1&#8221; and &#8220;2&#8221; indicate the level of occupancy detected.<\/span><\/p>\n<p><span style=\"color: #0000ff;font-family: Arial, helvetica, sans-serif;font-size: small\">To accommodate this data type&#8217;s use of &#8220;2&#8221;, the other <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/occupancy_estimation.htm\">occupancy models<\/a>, including the <a href=\"http:\/\/warnercnr.colostate.edu\/~gwhite\/mark\/markhelp\/occupancy_estimation_royle_nichols.htm\">Royle\/Nichols occupancy models<\/a>, treat the &#8220;2&#8221; encounter code as a &#8220;1&#8221;, and do not distinguish between them as does this data type.<\/span><\/p>\n","protected":false},"excerpt":{"rendered":"<p>Contents &#8211; Index Occupancy Estimation Multiple States Single Season Estimation of occupancy rates generally assume that a site or plot is occupied or not occupied.\u00a0 This model extends this concept to not occupied, and 2 levels of occupancy.\u00a0 For example, &hellip;<\/p>\n<p class=\"read-more\"> <a class=\"more-link\" href=\"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/occupancy-estimation-multiple-states-single-season\/\"> <span class=\"screen-reader-text\">Occupancy Estimation Multiple States Single Season<\/span> Read More &raquo;<\/a><\/p>\n","protected":false},"author":117,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-280","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages\/280","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/users\/117"}],"replies":[{"embeddable":true,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/comments?post=280"}],"version-history":[{"count":1,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages\/280\/revisions"}],"predecessor-version":[{"id":281,"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/pages\/280\/revisions\/281"}],"wp:attachment":[{"href":"https:\/\/sites.warnercnr.colostate.edu\/gwhite\/wp-json\/wp\/v2\/media?parent=280"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}