Ph.D. Biology, University of California, Santa Cruz, December 1999
Dissertation: Mammalian carnivores, mesopredator release, and avifaunal extinctions in a fragmented system.
Advisor: Michael Soulé, Department of Environmental Studies
M.S. Ecology, University of California, Davis, May 1994
Thesis: Comparative ecology of the island spotted skunk and the island fox of Santa Cruz Island, California.
Advisor: Dirk Van Vuren, Department of Wildlife, Fish and Conservation Biology
B.S. Zoology (Honors), Colorado State University, May 1989
Honor’s Thesis: The relationship between kinship and affiliative behavior in a free-ranging troop of primates.